Key to the subspecies
1a. Leaves strigose to slightly pilose below; branches covered with scalesa. ssp. malabathricum
b. Leaves pilose beneath; branches piloseb. ssp. normale
- Protologue
- Synonyms
- Vernacular names
- Description
- Distribution, ecology and notes
- Collections

Melastoma malabathricum L., Sp. Pl. 1 (1753): 390. - Type: Wight s.n., India orientalis (LINN, not seen).
foliis lanceolato-ovatis quincenervibus scabris quincenervia hirta major, capitulis fericeis villosis.

- Melastoma affine D. Don, Mem. Wern. Nat. Hist. Soc. 4 (1823): 288. - Type: Roxburgh s.n., India orientalis, (K?, not seen).
- Melastoma obvolutum Jack, Trans. Linn. Soc. London, Bot. 14 (1823): 3. - Melastoma malabathricum var. obvolutum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 80. - Type: Korthals s.n., Padang besie, Sumatera Barat, Indonesia (L, not seen).
- Melastoma denticulatum Labill., Sert. Austro-Caled. 2 (1824): 65, t. 64.
- Melastoma asperum Blume, Bijdr. 17 (1826): 1076. - Melastoma trachyphyllum (Blume) Backer ex Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 85. - Type: Blume s.n., Java, Indonesia (K!, P!).
- Melastoma sylvaticum Blume, Bijdr. 17 (1826): 1077. - Type: Blume s.n., Java, Indonesia (K!, L!).
- Melastoma taitense DC., Prodr. 3 (1828): 144. - Type: Gaudichaud s.n., Tahiti, Society Islands (P!).
- Melastoma polyanthum Blume, Flora 14 (1831): 481. - Type: Blume s.n., Baatavia, Jawa Barat, Indonesia (K!, P!).
- Melastoma polyanthum var. pallens Blume, Flora 14 (1831): 482. - Type: Blume s.n., Bantam, Jawa Barat, Indonesia (L!).
- Melastoma polyanthum var. riparium Blume, Flora 14 (1831): 482. - Type: Blume s.n., ad ripas fluminis circa Buitenzorg, Jawa Barat, Indonesia (L!).
- Melastoma tidorense Blume, Flora 14 (1831): 482. - Type: Reinwardt s.n., Tidore, Maluku, Indonesia (L, not seen).
- Melastoma royenii Blume, Flora 14 (1831): 483. - Melastoma polyanthum var. royenii (Blume) Miq. - Type: van Royen s.n., Sri Lanka (C!).
- Melastoma decemdentatum Kostel., Allg. Med.-Pharm. Fl. 4 (1835): 1511. - Type: not designated.
- Melastoma jackianum Korth., Verh. Nat. Gesch. Ned. Bezitt., Bot. 7 (1844): 227. - Type: Korthals s.n., Padang Highlands, Sumatera Barat, Indonesia (L, not seen).
- Melastoma punctatum Korth., Verh. Nat. Gesch. Ned. Bezitt., Bot. 7 (1844): 229. - Type: Korthals s.n., G. Boerangrang, Jawa Barat, Indonesia, 1100 m (L, not seen).
- Melastoma pusillum Blume, Mus. Bot. Lugd. Bat. 1 (1849): 54. - Type: Korthals s.n., Pulau Lampé, Borneo, Indonesia (K!, P!).
- Melastoma tondanense Blume, Mus. Bot. Lugd. Bat. 1 (1849): 54. - Type: Reinwardt s.n., Celebes: prope Tondano, Sulawesi Utara, Indonesia, Oct 1821 (K!, P!).
- Melastoma vitiense Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 275. - Type: le Guillou s.n., Balaou, Fiji Islands (P!).
- Melastoma ceramense Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 276. - Type: Jaquinot s.n., Warou, Ceram, Maluku, Indonesia (P!).
- Melastoma mariannum Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 276. - Melastoma malabathricum var. mariannum (Naudin) Fosberg & Sachet, Smithsonian Contr. Bot. 45 (1980): 15. - Type: Hombron s.n., Guham-Archipel, Marianne Islands (P!).
- Melastoma anoplanthum Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 277. - Type: Gaudichaud s.n., Pulau Penang, Malaysia, Mar 1837 (P!).
- Melastoma hombronianum Naudin, Ann. Sci. Nat, Ser. 3, 13 (1850): 278. - Type: Hombron s.n., Sanboangon Point sudest de la presquile sudest de Mindanao, Zamboango del Sur, Philippines (P!).
- Melastoma paleaceum Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 279. - Type: Gaudichaud 111, Tourane, Quang Nam-Da Nang, Vietnam, Jan 1837 (P!).
- Melastoma pelagicum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 279. - Type: Hombron s.n., Isle Ysabel, Solomon Islands (P!).
- Melastoma articulatum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 285. - Type: Hooker s.n., India orintalis (P, not seen).
- Melastoma heterostegium Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 286. - Type: Callery s.n., Pangasinan, Albay, Philippines, 1840 (P!).
- Melastoma homostegium Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 286. - Type: Cuming 927, Manila, Rizal, Luzon, Philippines, 1841 (BM!, K!, P!).
- Melastoma fasciculare Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 288. - Type: Gaudichaud 294, Manila, Rizal, Manila, Nov 1836 (P!).
- Melastoma baumianum Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 289. - Type: Baume s.n., Great lake at Manila, Rizal, Luzon, Philippines, Aug 1832 (K!, P!).
- Melastoma longiflorum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 290. - Type: Gaudichaud 108, Tourane, Vietnam, Jan 1837 (P!)
- Melastoma novae-hollandiae Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 290. - Type: Verreaux 571; Moreton Bay, Neu-Holland, Queensland, Australia, Dec 1845 (P!).
- Melastoma sechellarum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 290. - Type: Pervillé s.n., Seychelles, 1841 (K!, P!).
- Melastoma ellipticum Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 291. - Type: Leschenault s.n., Sri Lanka (P!).
- Melastoma brachyodon Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 292. - Type: Zollinger 5, Java, Indonesia, May 1842 (P!).
- Melastoma oliganthum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 292. - Type: Gaudichaud 78 ter, Singapore, Feb 1837 (P!).
- Melastoma longifolium Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 293. - Type: Guillou s.n., Singapore, Jul 1839 (P!).
- Melastoma triflorum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 293. - Type: Maie s.n., Goa, India (P!).
- Melastoma trachycaulon Miq., Fl. Ind. Bat., Suppl. 1 (1860): 316. - Type: H. B. 826, Pulau Pisang, Padang, Sumatera Barat, Indonesia, H.B. 826 (U!).
- Melastoma pusillum var. longifolium Cogn., Monogr. Phan 7 (1891): 354. - Melastoma borneense Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 84 - Type: de Vriese 168, Borneo, Indonesia, 1860 (L!).
- Melastoma francavillanum Cogn., Monogr. Phan. 7 (1891): 358. - Type: Zollinger 2175, Mt. Waliran, Java, Indonesia, 2000 m, 28 Aug 1844 (P!).
- Melastoma fuscum Merr., Philipp. Gov. Lab. Bur. Bull. 17 (1904): 39. - Type: Barnes FB 340, Lamao River, Luzon, Bataan, Philippines, Feb 1904 (BM!, K!).
- Melastoma parvifolium Merr., Philipp. Gov. Lab. Bur. Bull. 29 (1905): 32. - Type: Elmer 5836, Benguet Province, Baguio, Luzon, Philippines, Mar 1904 (P!).
- Melastoma lanaense Merr., Philipp. J. Sci., Bot. 3 (1908): 153. - Type: Clemens 836, Lake Lanao, Camp Keithley, Lanao del Sur, Mindanao, Philippines, Nov 1906 (PNH?, not seen).
- Melastoma intermedium Dunn, J. Linn. Soc., Bot., London 38 (1908): 360. - Type: Dunn 2706, Yenping Mountains, Fukien, China, 1500 m, Dunn 2706 (P!).
- Melastoma membranaceum Merr., Philipp. Journ. Sci., Bot. 3 (1909): 423. - Type: Fénix BS 3798, Batanes Islands, Cagayan, Philippines, Jun 1907 (K!).
- Melastoma warrineri C.B. Robin., Philipp. J. Sci., Bot. 6 (1911): 352. - Type: Robinson BS 9370, Infanta, Quezon, Luzon, Philippines, 900 m, Aug 1909 (K!).
- Melastoma congesta Elmer, Leafl. Philipp. Bot. 4 (1911): 1194. - Type: Elmer 12402, Magallanes (Mt. Giting-Giting), Sibuyan, Capiz, Philippines, May 1910 (BM!, K!, L!).
- Melastoma scabrum Ridl., J. Straits Branch Roy. Asiat. Soc. 79 (1918): 66. - Type: Coll. ign. 150113, Selaya Tupok, Burau, Langkawi, Kedah, Malaysia, Apr 1911 (K!).
- Melastoma holmani Elmer, Leafl. Philipp. Bot. 8 (1919): 3089. - Type: Elmer 18479, Los Baños (Mt. Maquiling), Laguna, Luzon, Philippines, June-July 1917 (PNH?, not seen).
- Melastoma vulcanicum Ridl., J. Malayan Branch Roy. Asiat. Soc. 1 (1923): 60. - Type: Ridley s.n., Sibayak Volcano, Berastagi, Bigkeerk (?), Sumatera Utara, Indonesia (K!).
- Melastoma polyanthum var. pulleana Mansf., Engl. Bot. Jahrb. 60 (1924): 109. - Type: Pulle 554, Perameles Mts., Papua New Guinea, 1000 m (L, not seen).
- Melastoma subgrande Hochr., Candollea 2 (1925): 486. - Type: Hochreuter 3387, Île dUpolu, lac Lanunea, Samoa, 500 m, 25. Mar 1905 (G?, not seen).
- Melastoma roemeri Mansf., Engl. Bot. Jahrb. 60 (1926): 107. - Types: von Römer 1123, Hellwig-Gebirge, Madang, Papua New Guinea, 1000-1500 m, Nov 1909 (L!); Janowsky 190a, Siriwo-Fluß , Irian Jaya, Indonesia, Jul 1912 (L!); von Römer 395, Lorentzfluß , Irian Jaya, Indonesia, 2. Oct 1909 (L!); Versteeg 1598, Lorentzfluß bei Alkmaarbivak, Irian Jaya, Indonesia, 11. Aug 1907, 150 m (L!).
- Melastoma imbricatum var. longipes Craib, Fl. Siam. 1 (1931): 679. - Type: Put 224, Baw Re, Chantaburi, Thailand, 22 Jul 1926 (K!).
- Melastoma malabathricum var. grandiflorum Craib, Fl. Siam. 1 (1931): 681. - Type: Kerr 15235, Khao Sung, Phattalung, Thailand, 900 m, 10 Apr 1928 (BK!, BM!).
- Melastoma pinatubense Elmer, Leafl. Philipp. Bot. 9 (1934): 3204. - Type: Elmer 22233, Camp Stotsenburg (Mt. Pinatubo), Pampanga, Luzon, Philippines, May 1927 (SING, K!, C!, BM!, P!, L).
- Melastoma stenophyllum Merr., Mich. Acad. Sci. Arts Letters 24 (1938): 83. - Type: Rahmat 7000, near Hoeta Bagasan, Sumatra, Indonesia, 7 Sep 1934 to 4 Feb 1935 (PNH?, not seen).
- Melastoma polyanthum var. linearifolium Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 69. - Type: not designated.
- Melastoma polyanthum var. mollissimum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 70. - Type: Toxopeus 217, Nal Besi, Boeroe, Maluku, 14 Jun 1921, 950 m (L!).
- Melastoma suave Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 74. - Type: Teysmann s.n., Java, Bogor, Jawa Barat, Indonesia, Jul 1867 (L!).
- Melastoma malabathricum var. javanum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 82. - Type: Elbert 192, Lawoe, Res. Madioen, Jawa Timur, Indonesia, 1400 m, 14 Sep 1912 (L!).
- Melastoma sylvaticum var. permultiflorum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 84. - Type: Lörzing s.n., Karohaugrlakten (?) aan de Sibayak, Sumatera Utara, Indonesia, 1450 m, 25 May 1921 (L!).
- Melastoma trachyphyllum var. ochraceum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 87. - Type: Lörzing 5995, southeast of G. Sibajak, Sibaulangit, Sumatra, Indonesia, 1700-2000 m., 28 Aug 1918 (L!, P!).
- Melastoma caesium Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 88. - Type: Elbert 4187, north side of Batu-Lanteh Mtns, Sumbawa, Nusa Tenggara Barat, Indonesia, 1700 m, 20 Jan 1910 (L!).
- Melastoma buennemeyeri Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 91. - Type: Bünnemeyer 881, Talakmau), Sumatera Barat, Indonesia 1900 m, 27 May 1917 (L!).
- Melastoma robustum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 92. - Type: Bünnemeyer 6585, G. Walk., P. Lingga, Lingga Archipelago, Riau, Indonesia, 300 m, 12 Jul 1919 (L!).
- Melastoma patulisetum Ohwi, Bot. Mag. Tokyo 57 (1943): 1. - Type: Kanehira-Hatusima 13180, Ware, 60 miles south of Manokwari, Irian Jaya, Indonesia, 500 m, 26 Mar 1940 (TNS?, not seen).

Vernacular names: Harendong (Java); Sikadoekdoek (Sumatra); Tjengkodok (Borneo); Gosing (D/Bundu Tuhan, Borneo); Sendudok/Sendudok Gunong, Kuduk-kuduk (Malaysia); Aiinot (Enga, Yogos, New Guinea); Djuk (Yoowi dialect, Hagen-Chimbu language, New Guinea); Davun-dava (Solomon Islands); Camoteng/Kamutin (Tagbanua, Philippines); Bayakin-kin (Mang, Philippines); Singapore Rhododendron/Straits Rhododendron (English); Maha-bowitiya (Singapore); Aa-noi Ton, Klong kleng, Beng-ah, Mai grá (Thailand).

Shrubs 1.5--3 m high or small trees up to 5 m tall; young branches quadrangular, covered with appressed or slightly spreading scales, bark grey or brown; old branches terete, glabrescent. Leaves elliptic to lanceolate, 6--15 x 2--6.5 cm, occasionally narrowly lanceolate to oblong, 4--6 x 0.6--1.5 cm, base rounded to acute, apex acuminate; lamina strigose above, beneath strigose to slightly pilose; nerves 5 or 7, beneath covered with appressed to spreading scales; petiole 7--20 mm long. Inflorescences terminal cymes of 3--12 flowers. Flowers 5-merous, seldom 6-, 7-, or 8-merous, often varying on a single plant; bracts deciduous, lanceolate or ovate, acuminate, varying from small and inconspicuous (2.3 x 2.0 mm) to large and enclosing the buds (16 x 14 mm), outside covered with small appressed scales at least along the midrib, inside glabrous. Hypanthium campanulate, varying from 5 x 5 mm to 11 x 10 mm, covered with appressed or slightly spreading, (0.8--)1.2--2.5(--3.4) mm long, golden to red scales; sepals lanceolate, (3.2--)4.5--7.0(--10.5) x (1.9--)2.5--3.5(--6.2) mm, outside covered with appressed scales at least along the midrib, inside glabrous or hairy at the apex; intersepalar emergences subulate, (1.0--)1.5--3.5(--5.5) mm long. Petals obovate, 15--35 x 10--22 mm, ciliate at the apical margin, violet, seldom white (especially on the Pacific Islands). Stamens dimorphic (seldom isomorphic); anthers of longer stamens 6.5--13.1 mm long, slightly sigmoid, violet, connectives 9.7--17 mm long, with a ventral bilobed, yellow appendage c. 0.9--3.3 mm long, filaments 6--12 mm long; anthers of shorter inner stamens 5.5--10.5 mm long, ± straight, yellow, connective not prolonged or seldom up to 2.0 mm long, with 2 ventral appendages c. 0.6--2.0 mm long, yellow, filaments 5.5--9.5 mm long. Ovary shorter than the hypanthium, crowned by golden bristles; style 23--32 mm long, sigmoid. Fruit a fleshy capsule, 6.5--10(--11.5) x (5--)6--9(--10.5) mm, rupturing irregularly transversally at maturity, exposing the soft, dark blue pulp with the orange seeds.

Distribution - Indian Ocean (Mauritius, Seychelles), South and Southeast Asia, South China, Taiwan, Australia, South Pacific Ocean (Micronesia, Melanesia, Polynesia).
Click to get a larger map. Map made with Online Map Creation.
Ecology - Collections: 338. In disturbed places (roadsides, river banks, secondary forests), on fallow land, or on grasslands from sea level to up to 2900 m altitude.
Notes - Melastoma malabathricum is the most widespread and morphologically most variable species of the genus, resulting in the description of many separate species. After examining numerous specimens from the entire distributional range, I found intermediate forms connecting the types of the named entities. Their characters combine nearly independently within the geographical range of M. malabathricum which makes it impossible to describe subspecies or varieties based on more than one or two collections. An exception is ssp. normale, which is more pilose than ssp. malabathricum and grows in the northern part of the distribution area. Intermediate forms in branch indumentum and specimens from the Malay Archipelago do not allow to treat ssp. normale as a separate species.
The great variability of M. malabathricum may be connected with its preferred habitat. The plants grow in disturbed places that today are more common in Asia and Oceania than undisturbed ones. It is possible that the species expanded its natural area synanthropically resulting in the isolation of small populations and the subsequent stabilization of morphological characters of the (self-compatible) founder individual.

AUSTRALIA.
- Helms 1189, North Queensland, Barren Falls near Cairns, Queensland (C)
- Hyland 8394, TR 14 (McIlwraith Range Leo Ck. Road), Queensland, 450 m (L)
- Kanis 2157, Cook Distr., Herberton State Forest, southwest of Atherton, Queensland (L)
- Specht 863, Arnhem Land Aboriginal Reserve, Yirrkala, Northern Territory (L)
- v. Mueller s.n., Rockingham Bay, Queensland (C)
BRUNEI
- Forman 825, Bkt. Beruang, south of Sg. Penyatang, Tutong, Brunei, 20 m (C, K, L)
- Wong 393, nothwest of Bt. Retak, knoll, Temburong, Brunei, 1260 m (SAN)
CAROLINE ISLANDS
- Stone 5417, Ponape
- Mt. Seletenreh, U District, trail ascending northwest face, through Campnosperma-Parinarium-Myristica forest, , Caroline Islands, 600 m (L)
CHINA
- Coll. ign. s.n., East China (C)
- Dunn 2706, Yenping Mountains, Fukien, 1500 m (P)
- Fung 20067, Paai Poon Tsuen and vicinity (Ngai District), Hainan, Hainan (P)
- Lei 636, Tim Neung Shu, Pak Shik Ling and vicinity, Ku Tung village, Ching Mai Distr., Hainan (L)
- Lin 11986, Nantai, Foochow, Nongli, Fukien (C)
- Tang 4161, Foochow and vicinity, near YMCA retreat, Fukien (C)
- Tang 5862, Kushan monastery, Gi-shan, Fujian (C)
- Tang 5948, Foochow and vicinity, North Park, Fukien (C)
- Tsang 26578, Na Leung and vicinity, Fang Cheng District, Guangxi (P, C)
FIJI ISLANDS
- Nielsen 282, Waidoi, Viti Levu (C)
- Smith 5439, Nandronga & Navosa, north portion of Rairaimatuku Plateau, between Nandrau and Rewassau, Viti Levu, 825 m (L)
FRANCE
- Baas-Becking 5586, St. Louis, New Caledonie (GOET)
- Croockewit s.n., Mt. Koghis, Nouvelle-Calédonie, 600 m (L)
- Didrichsen s.n., Tahiti, French Polynesia (C)
- Ostenfeld 184, Papeari, Tahiti, French Polynesia (C)
- Tobrié s.n., Society Islands (?), French Polynesia (C)
GUAM
INDIA
- Barnes 118, Travangore, Travancore and Cochin (K)
- Børgesen 194, Arbail Ghat between Kanrar and Dhasuar, Bombay (C)
- Coll. ign. 2875, Little Nicobar, Nicobar Islands (C)
- Comy 2244, East Bengal (C, GOET)
- Hohenacker s.n., Mangalore, prope urbem (C)
- Hooker s.n., Malabar, Concan (C, GOET)
- Hooker s.n., Sikkim, 1000 m (K)
- Hooker s.n., Mt. Khasia
- regio trop., Meghalaya (C, GOET)
- Kamphövener 2421, Teressa, Nicobar Islands (C)
- Kamphövener 2654, Kakana, Nicobar Islands (C)
- Kamphövener 2811, Ytoe, Nicobar Islands (C)
- Mooney 1474, Bailadila Hills, Bastar, 1000 m (K)
- Mueller s.n. (C)
- Müller s.n. (C)
- Parry 1111, Distr. Garo Hills, Dalu, Assam, 500 m (K)
- Saldanha 16288, Hassan Distr., along the ghat road, Charmadi ghat, Mysore (K)
- Stocks s.n., Maisor & Carnatic, Mysore (C)
- Voigt 132, Botanical Garden of Calcutta and Serampore, West Bengal (C)
- Voigt 1564, Botanic Gardens Calcutta and Serampore (C)
- Voigt s.n., Botanic Gardens Calcutta and Serampore, West Bengal (C)
INDONESIA
- Alston 15676, Tomohon, Sulawesi Utara, 700 m (L)
- Bakhuizen v. d. Brink 5657, Res. Preanger Reg.
- G. Perbakti (west of halte Tjitjoeroeg), Jawa Barat, (L)
- Bloembergen 4287, Island Soela Sanana doorstrek Sanana-Molboefa, Maluku, 100 m (L)
- Blume s.n., Baatavia, Jawa Barat (K, P)
- Blume s.n., Java (K)
- Blume s.n., Java (K)
- Blume s.n., Java, prope Artja (K, L)
- Blume s.n., Java (K, P)
- Blume s.n., Java, Jawa (C)
- Blume s.n.
- ad ripas fluminis circa Buitenzorg, Jawa Barat (L)
- Blume s.n.
- Bantam, Jawa Barat (L)
- Bünnemeyer 881, Sumatra nortwest coast, Talakmau (= G. Ophyr), Sumatera Barat, 1900 m (L)
- Bünnemeyer 6585, Lingga Archipelago, P. Lingga, G. Walk., Riau, 300 m (L)
- Bünnemeyer 9798, west coast
- G. Koerintji, Sumatera Barat, 2220 m (L)
- Cellinese 73, Barito Ulu, Rekut Camp, Kalimantan Tengah (BO)
- Coll. ign. s.n., Java (K)
- Coll. ign. s.n., Java (K)
- Coll. ign. s.n., Sumatra,(C)
- Elbert 192, Lawoe, Res. Madioen, Jawa Timur, 1400 m (L)
- Elbert 2151, Lombok, Rindjani Vulkangebirge, Südseite, Abhang oberhalb Reinbang-Rerang, Nusa Tenggara Barat, 1900 m (L)
- Elbert 4187, Sumbawa, north side of Batu-Lanteh Mtns, Nusa Tenggara Barat, 1700 m (L)
- Elmer 13059, Puerto Princesa (Mt. Pulgar), Palawan ()
- Enoh 266, Pontianak, Soemoer bor. Soengai bangkong, Kalimantan Barat, 3 m (SING, BO)
- Eyma 4174, Rec. Selebes & Onderhoorigheden subdiv. Malili, between Takolekadjoe & Tawi baroe, Sulawesi Selatan (L)
- Franck 167, southeast Borneo, Mardhapura (=Martapura), Kalimantan Selatan (C)
- Franck 168, Idjen Plateau, Java (C)
- Franck 1089, Vad G. Aujer get Jaoa, Jawa Timur (C)
- H.B. 826, Pulau Pisang, Padang (U)
- Hagerup s.n., Toba, Sumatra, 1200 m (C)
- Hansen 1256, Malacca: Johore, Kalimantan Tengah, 50 m (C)
- Hickson s.n., Indian archipelago, Koeang shore (= Air Muang-binjai, Sumatra?) (K)
- Jacobs 4416, south slope of Mt. Kerintji, path from Kaju Aro towards the top, Sumatera Barat, 2900 m (L)
- Jacobs 8193, Mt. Tanggamus, Sumatra, Lampung, 1400 m (C, KEP, L)
- Janowsky 190 a, flum. Sirirov, Irian Jaya (L)
- Jarvie 5032, Kalteng
- Samba. At about km 45, Kalimantan, 235 m (BO)
- Jarvie 6122, Kalteng
- Samba. Path down from summit of Bkt. Raya, Kalimantan, 2310 m (BO)
- Junghuhn s.n., Sumatra, Pulo Pontjang Pitpil at Tapanoelic Gai, Sumatera Utara (L)
- Korthals s.n., Borneo (C)
- Korthals s.n., Borneo (C)
- Korthals s.n., Java (C)
- Korthals s.n. (L)
- Kostermans 18802, West Sumbawa, northwest slope of Mt. Batulanteh, Nusa Tenggara Barat, 500 m (L)
- Kostermans 19176, West Sumbawa, Mt. Batulanto, ridge to top, Nusa Tengara Barat, 1300 m (C, L, P)
- Kostermans 430, Bangka, Lobok-besar, Sumatera Selatan, 20 m (L)
- Lörzing 13050, Medan, Sumatera Utara, 20 m (L)
- Lörzing s.n., Sumatra, Karohaugrlakten aan de Sibayak, Sumatera Utara, 1450 m (K, L)
- Maskuri 6, West Java
- Poreang. Mt. Pameungpeuk - South Sukabumi, Jawa Barat, 1200 m (L)
- Nielsen 1079, Emmahafen, Sumatra (C)
- Nielsen 1300, Vulkan Pintau, Sumatera Utera (C)
- Nielsen 1318, Vulcan Pintau, Sumatera Utera (C)
- Reinwardt s.n., Java (C)
- Reinwardt s.n., prope Tondano, Sulawesi Utara (K, P)
- Ridley s.n., Sibayak Volcano, Berastagi, Bigkeerk?, Sumatera Utara (K)
- Ridsdale PBU 380, Project Barito Ulu base camp and environs, Kalimantan Tengah, 0 m (L)
- Robinson 40a, Korinchi Peak, Sumatera Barat, 2190 m (BM)
- Robinson 40b, Siolak Dras, Korinchi, Sumatera Barat, 1000 m (BM)
- Robinson 40c, Korinchi, Sumatera Barat, 2200 m (B)
- Royen 3208, Radjah Ampat Distr., Klabala river southeast of Sorong, Irian Jaya, 35 m (L)
- Royen 3995, District Manokwari, Kebar valley, c. 100 km west of Manokwari, between Andjai Village and Andjai Airstrip, Irian Jaya, 550 m (L)
- Schmutz 1808, West-Ruteng, Potjo Rii-Potjo Ngando Napu, Flores, Nusa Tenggara Timur, 1700 m (L)
- Symington FMS 23985, Barastagi, Sumatra (KEP)
- Teysmann s.n., Java, Bogor (L)
- Thiv s.n., road Ciawi-Cirurug, 5-10 km east of road, Jawa Barat, 600 m (MJG)
- Versteeg 1598, flum. Lorentz, Alkmaarbivak, Irian Jaya, 150 m (L)
- Vogel 5701, Soroako, south shore of Lake Matano, Sulawesi Selatan, 400 m (L)
- Westermann s.n., Java (C)
- Yates 1976, East Coast, Sumatra (SING, BO)
MALAYSIA
- Amin SAN 96908, Kretam Forest Reserve, Sandakan Distr., Sabah (SAN, PSU)
- Awa S 46908, Dulit Range, Tinjar, 4th Division, Sarawak, 800 m (SAR,AAU,C,L,K,KEP)
- Beccari 3974, Sarawak (K, P)
- Brooke 8000, Kuching, 1st Div., Sarawak (BM)
- Burkill HMB 281, Pangkor Island, Pangkor, Perak, 7 m (SING, L)
- Burkill 12622, Birchs Hill, Perak, 1300 m (SING)
- Clausing 19, Poring Hot Springs, oberes Tor an der Straße, Sabah (Mt. Kinabalu Herbarium)
- Clausing 102, Kinabalu National Park, East Mesilau River, Pinosuk Plateau, Sabah, 1800 m (KEP)
- Clausing 155, Trail to Paku Beach, Bako National Park, Sarawak (UNIMAS, MJG)
- Clausing 187, Bukit Matang, Sarawak (UNIMAS)
- Clausing 213, Tenompok, Mt. Kinabalu National Park, Sabah, 1600 m (Mt. Kinabalu Herbarium)
- Clausing 242, Mammut Copper Mine, Ranau, Sabah (Mt. Kinabalu Herbarium)
- Coll. ign. 52, Pulau Penang (BM)
- Coll. ign. 150113, Selaya Tupok, Burau, Langkawi, Kedah (K)
- Corner 33167, Frasers Hill, Pahang, 1200 m (SING)
- Corner s.n., Sg. Tahan, Pahang (SING)
- Corner s.n., Sg. Tahan, Pahang (SING)
- Cuadra A 2267, Sg. Sapaganga, Elopura, Sandakan, Sabah (SING, KEP)
- Davis 69208, Genting Highlands, G. Ulu Kali, summit area, Pahang, 1700 m ()
- Dewol SAN 67378, Pinangah FR, Pinangah Distr., Sabah, 800 m (SAN, CMU, K, L)
- Didrichsen 3573, Penang, Penang (C)
- Elmer 20039, Sandakan
- Myburgh Province, Sabah (C, BM)
- Everett FRI 13578, Taiping: 3,25 mile up road. Road verge hill side, over acidic rock, Perak, 680 m (SING, KEP)
- Everett FRI 13644, Central Pahang, low undulating country by logging track, Pahang, 90 m (SAN, KEP)
- Hansen 231, Gunong Mulu National Park, camp 5 close to Helipad, Sarawak, 150 m (C)
- Hansen 1008, G. Serapi, 1st Div., Sarawak, 600 m (C)
- Ilias S 42541, path to G. Silantek, Ulu Sg. Silantek, Kiri, 85th mile Simanggang Road, 2nd Division, Sarawak, 216 m (SAR, C, L, K, SAN, MO)
- Ilias S 43789, Niah Forest Reserve, 4th Division, Sarawak, (SAR, AAU, C, L, K, KEP)
- Kochummen FRI 18408, Kg. Jawa Serting Tengah Valley, Negeri Sembilan (KEP)
- Lee S 44512, Bt. Tangga Saran, Engkilili, 2nd division, Sarawak, 830 m (SAR, AAU, L, K, KEP)
- Lee S 45343, Sg. Iban, Linau, Belaga, 7th Div., Sarawak (SAR, K, L, KEP, SAN, AAU)
- Lee SAN 96893, 5 miles from Kampung Mendulong road to Meligan, Sipitang distr., Sabah (SING, SAN)
- Mat-Salleh 3402, Telupid, Bt. Tawai ForestReserve, south trail, along Sg. Meliau, Sabah, 100 m (SAN)
- Maxwell 78158, Cameron Highlands
- G. Brinchang, Pahang, 1900 m (L)
- Meyer 9518, G. Penrissen, near Indonesian border, access road, Sarawak, 600 m (KEP)
- Meyer 9519, G. Penrissen, near Indonesian border, Sarawak, 1000 m (KEP)
- Meyer 9520, G. Penrissen, near Indonesian border, Sarawak, 1000 m (KEP)
- Meyer 9531, Kinabalu National Park, Headquarter, end of Liwagu River Trail at Ring Road, Sabah, 1600 m (KEP)
- Meyer 9532, Kinabalu National Park, Liwagu River Trail near Headquarter, Sabah, 1500 m (KEP)
- Meyer 9533, Kinabalu National Park, Summit Trail at km 0.5, Sabah, 2000 m (KEP)
- Meyer 9534, Kinabalu National Park, Powerstation, Sabah, 1900 m (KEP)
- Meyer 9536, Kinabalu National Park, Headquarter, near staff quarters, Sabah, 1550 m (KEP)
- Meyer 9538, Kinabalu National Park: Poring Hot Springs, road near Park Entrance, Sabah (KEP)
- Meyer 9539, Kinabalu National Park: Poring Hot Springs, road near Park Entrance, Sabah (KEP)
- Meyer 9540, Sandakan: Labuk Road mile 14, Sabah (KE)
- Meyer 9541, Sandakan: side road of Labuk Road mile 14 (Jln Fook Hin), Sabah (KEP)
- Meyer 9550, Cameron Highlands: road to G. Brinchang above Sungai Palas Tea Estate, Pahang, 1800 m (KEP)
- Meyer 9551, Cameron Highlands: G. Brinchang, near summit, Pahang, 2000 m (KEP)
- Meyer 9553, Cameron Highlands: Tanah Rata roaad, ca. 1 km above Habu Powerstation, Pahang, 1300 m (KEP)
- Meyer 9554, Cameron Highlands: Tanah Rata road, side road at mile 33, Pahang, 1400 m (KEP)
- Meyer 9637, Taiping, road Taman Tasik-Bt. Larut, ca. 100 m to entrance to Bt. Larut, Perak, 100 m (KEP)
- Meyer 9639, Kota Bharu, 5.75 mile PCB road, Kelantan, 10 m (KEP)
- Meyer 9641, Pulau Tioman, Kp. Tekek, ca. 100 m above mosque on trail to Kp. Juara, Pahang, 50 m (KEP)
- Ng FRI 27026, Sg. Tahan, Pahang (KEP, L)
- Pawanche FMS 13422, Bt. Beruang, Kepong Selangor, Selangor (SING, KEP)
- Puni 17, Penang, near RECSAM, Glugor, Penang, 25 m (L)
- Ridley 16036, G. Tahan, Pahang (SING)
- Sato 1148, Tamparuli, Daerah, Tuaran, Sabah, 150 m (SAN)
- Seimund s.n., Kuala Teka, Pahang (K)
- Shah MS 1105, Maxwells Hill: summit of G. Hijau, Perak, 1350 m (SING, C, L)
- Shah MS 1118, Maxwells Hill, 4th mile, Taiping, Perak, 600 m (SING, L)
- Shah MS 1923, Ulu Sg. Sepia near Kuala Aur, Pahang, 45 m (SING, KEP, C)
- Shah MS 2671, Taman Negara: Sg. Tahan, Pahang (SING, KEP, C)
- Smith FRI 63659, Cameron Highlands, Mentiji Plantation, Pahang (KEP)
- Stone SAN 87059, Pulau Balembangan, northside of Periok headland, Kudat Distr., Sabah, 1 m (SAN)
- Symington FMS 33209, Klang Gater, Selangor (SING)
- Vesterdal 401, Malacca, Johore (C)
- Vesterdal 513, Malacca (C)
- Vethevelu FRI 32903, Genting Highlands, Pahang, 1200 m (SAN, KEP, L)
- Wallich 4041, Penang (K)
- Warwick MW 51, Logging area about 20 m east of Telupid, Gunong Monkobo, Crocker Range near Telupid, Sabah, 1200 m (SAN, E, KEP)
- Whitmore FRI 3934, Dungun, Bt. Bauk Forest Reserve. Compt 20B mile 10 Bt. Besi Rd., Terengganu (SING, KEP)
- Whitmore FRI 12976, Langkawi Island, south coast 4 m. west of Kuah, Kedah (SING, KEP)
- Whitmore FRI 15398, banks of Sg. Tembeling between J. Fanjang and K. Tahan, Pahang (SAR,KEP)
- Wight 1105, Peninsula Indiae Orientalis ()
- Wray 5514, G. Tahan, Pahang, 1500 m (BM)
- Yii S 40687, Semengoh Forest Reserve, Biological Centre, Kuching, 1st Div., Sarawak, 30 m (KEP, SAR, UA, K, L, C)
- Zainudin 4857, Sandakan
- Telupid, Sungai Meliau off Sungai Karamuk, south trail, Sabah, 100 m (SAN)
MAURITIUS
- Coll. ign. 91, Mauritius (K).
NEPAL
- Wallich 4039b, Napalia (K)
PAPUA NEW GUINEA
- Brass 24672, Goodenough Island, east slopes, Milne Bay, 1450 m (L)
- Conn 1494, Manki Divide, southwest of Bulolo, Morobe, 1700 m (L)
- Craven 1194, near Haumga, Aseki Patrol Area, Morobe, 1400 m (L)
- Croft NGF 13172, Mt. Sule, about 25 miles northnortheast of Fulleborn Harbour, Subdistr. Pomio, East New Britain, 1500 m (L)
- Dissing 1996, Bismarck Archipelago (C)
- Dissing 2330, Bismarck Archipelago (C)
- Dissing 2607, New Britain, Cape Hoskins: deb af plt. fra vulkan awrodet 5 km ost fra Mount Bauco, Bismarck Archipelago (C)
- Foreman LAE 60417, Mear Woroi village on Oriomo River, Daru Subdistrict, Western, 20 m (L)
- Hartley 10263, near the Butibum River about 7 miles north of Lae, T.N.G, Morobe, 60 m (C)
- Hoogland 6753, near Poio village, Yaki valley, Wabag subdistr., Western Highlands, 2100 m (L)
- Køie 1525, Mussau, Bismarck Archipelago (C)
- Køie 1704, Mussau, Malakata, Bismarck Archipelago, 15 m (C)
- Millar NGF 35164, Amboin, Angoram Subdistrict, East Sepik, 105 m (L)
- Ridsdale NGF 33612, Near Weam, Bensbach Subdistr., Papua, Western, 30 m (SING, L, BRI, CANB, A, K, BO, SYD)
- Royen NGF 18241, confluent of Warapuri and Warrangga Rivers, Wahgi-Jimmy Divide, N. of Nondugl, Minj subdistr.,Eastern Highlands, 2100 m (SING, L)
- Sands 1432, Kikori Subprovince
- c. 213 km upstream from the coast on the Purari River
- near the river 5 km upstream from the Pikae tributary river and 13 km from Wabo, Gulf Province, 125 m (C)
- Saunders 50, Cape Nelson, lower slopes of Mt. Trafalgar northeast Tufi Subdistr., Northern, 480 m (L)
- Streimann NGF 44314, Lasanga Ils., Lae subdistr., Morobe, 3 m (L)
- v. Römer 1123, in montosis Hellwig, Madang, 1600 m (L)
- Vandenberg NGF 42114, shore of Lake Kopiago, Western Highlands, 1260 m (L)
- Vink 16424, Western Highlands
- Kubor Range: Uinba, Nona-Minj Divide, Western Highlands, 2000 m (C, L)
PHILIPPINES
- Barbon PPI 8313, Mt. Isarog, Naga City, Camarines Sur (BRIT)
- Barnes FB 204, Lamao River, Luzon, Bataan (K)
- Barnes FB 340, Lamao River, Luzon, Bataan (K, BM)
- Borden FB 2100, Lamao River, Mt. Mariveles, Luzon, Bataan (K)
- Borden FB 2709, Lamao River, Mt. Mariveles, Luzon, Bataan (K, SING)
- Broada 1380, North Luzon, Benguet (K)
- Castro PNH 6486, Anibawan, Polillo Island, Quezon (L)
- Copeland FB 298, Lamao River, Luzon, Bataan (K)
- Cordero PNH 91514, Lipuun, Quezon, Palawan (SAN, PNH)
- Cuming 927, Manila, Luzon, Rizal (K, BM, P, L)
- Edaño BS 79355, Mt. Mapolapola, Camiguin Island, Babuyan Islands, Cagayan (SING)
- Edaño PNH 1109, Mt. McKinley, east slope, Davao, 1900 m (L)
- Edaño PNH 34481, Mt. Malinao, Albay, 460 m (L)
- Elmer 9481, Dumaguete (Cuernos Mts.), Negros, Negros Oriental (BM)
- Elmer 12402, Magallanes (Mt. Giting-Giting), Sibuyan, Capiz (K, BM, L, MO)
- Elmer 14006, Cabadbaran (Mt. Urdaneta), Mindanao, Agusan del Norte (BM, L, MO)
- Elmer 15083, Irosin (Mt. Bulusan), Luzon, Sorsogon (C)
- Elmer 15452, Irosin (Mt. Bulusan), Luzon, Sorsogon (C)
- Elmer 17623, Los Baños (Mt. Maquiling), Luzon, Laguna (C)
- Elmer 21930, Camp Stotsenburg (Mt. Pinatubo), Luzon, Zambales (SING, P, C, L)
- Elmer 22233, Camp Stotsenburg (Mt. Pinatubo), Luzon, Pampanga (SING, K, C, BM, P, L)
- Gaerlan PPI 11009, Alvenda, Mutia, along the road, Zamboango del Norte (BRIT)
- Gates 5194, Mt. Maquiling, Laguna, 1200 m (BRIT)
- Gaudichaud 48, Manila, Rizal (K, P)
- Gaudichaud 294, Manila, Rizal (P)
- Iwatsuki P 614, en route from Mt. Bubonbilit to miners camp in Mt. Sicapao range, Ilocos Norte, 1900 m (L)
- Maliwanag 308, Bongabon and Pinamalyan, Mindoro, Pinamalayan (BRI)
- Merrill 7214, Palawan (K, P, L)
- Merrill BS 9645, Luzon, Benguet (L)
- Merrill FB 2104, Subic and vicinity, Luzon, Zambales (K)
- Ramos BS 44732, Mt. Tapolao, Luzon, Zambales (L)
- Ramos BS 47236, San Mariano, Luzon, Isabela (C)
- Reynoso PPI 7368, Visayas Isl, Sitio Kaguyanan, Brgy. Lokilokon, Paranas, along Kaguyanan Creek, Western Samar (BRIT)
- Reynoso PNH 87713, Mono, Quezon (SAN, PNH, L)
- Robinson BS 9370, Infanta, Luzon, Quezon, 900 m (K)
- Sandermann Olsen 530, Balabac: Dalawan Bay, Palawan, 5 m (C)
- Sandermann Olsen 665, Tarawakan, Tawi-Tawi, Sulu Archipelago, Sulu, 50 m (C)
- Sandermann Olsen 982, Sapamoro, 70 km northeast of Zamboanga, St. Crus. Isl., Mindanao, Zamboanga del Sur, 300 m (C)
- Serviñas BS 16904, Mt. Maquiling, Luzon, Laguna (BM)
- Sulit PNH 9901, Mt. Katanglad, south slope & east peak, Bo Kaatoan, Bukidnon, 1800 m (L)
- Whitford FB 38, Lamao River, Mt. Mariveles, Luzon, Bataan (K). SAMOA. Whitmee s.n., Samoa (C)
SEYCHELLES
- Jeffrey 666, Cascade, south side, Mahé, 540 m (L)
- Pervillé s.n., Seychelles (K, P)
SINGAPORE
- Bels 223, Alexandra Road (C)
- Salleh S 90, Pulau Senang,(C)
- Shah 1234, Botanic Gardens Singapore: Lawn C (E, SING)
SOLOMON ISLANDS
- Kajewski 2510, Guadalcanal, Vulolo, Tutuve Mtn, Solomon Islands, 1200 m (L)
- Tay 940071, Santa Cruz, Forestry Plantation, Solomon Islands, (SING, KEP)
SRI LANKA
- Coll. ign. s.n., Zeylanica (C)
- Jacobsen 13-10, Kosgoda (C)
- Msiha 23, Kalugrammane Distr., 750 m (C, BM)
- Robyns 7218, between Ulapane and Nawalapitiya, Kandy Distr., Central (C)
- Royen s.n., e Ceylona (C)
THAILAND
- Beusekom 1165, Ob Luang table land, along rd. from Bo Luang to Om Koi, Chiang Mai (AAU, BKF, C, E, K, KYO, L)
- Bunyaratappand 141, Phu Langka, Chiang Rai, 1290 m (BKF, C)
- Geesink 4998, Khlong Nang Yon, Phangnga, 100 m (AAU, BKF, C, E, P)
- Hansen 11870, Khao Phra Mi, foothills, Ranong, 70 m (BKF, C)
- Henningsen s.n., Singora, Songkhla (C)
- Hosseus 459, Doi Suthep National Park: Doi Suthep, c. 300-1400 m, Chiang Mai, 1000 m (E, C, P)
- Iwatsuki 8448, Khao Luang, middle elevation, Nakhon Si Thammarat, 750 m (C, KYO)
- Kerr 15235, Khao Sung, Phatthalung, 900 m (AAU, BK, E, BM)
- Kerr 19145, Khao Soi Dao, Trang, 400 m (BK, C, K)
- King 5466, Chiang Mai, 20 km west of, Chiang Mai, 1400 m (C)
- Larsen 798, 7 km east of Tung Salaeng Kuang forest station along trail, Phitsanulok, 600 m (AAU, BKF, C, K)
- Larsen 1435, 35 km north of Chumphon, Surat Thani, 50 m (AAU, C)
- Larsen 44331, Amphoe Tha Wang Pha: Doi Tiu, Nan, 1000 m (AAU)
- Maxwell 73325, Klong Mah Yom, Koh Chang, Trat, 50 m (AAU, BK)
- Meyer 956, Hat Yai: Ko Hong Hill, road to summit, Songkhla, 100 m (BKF)
- Meyer 959, Hat Yai: Prince of Songkhla University, Kuan Mod Dang Hill, in front of the guest house, Songkhla, (BKF)
- Meyer 9513, Khlong Hoi Khong Research Station, south of Hat Yai, Songkhla (BKF)
- Meyer 9514, Khlong Hoi Khong Research Station, south of Hat Yai, Songkhla (BKF)
- Meyer 9516, Khao Chong: west of Ban Na Wong, road to Trang, Trang, 150 m (BKF)
- Meyer 9618, Khao Khitchakut Nationalpark: near entrance on side road of Nationalroad 317, Chantaburi, 200 m (BKF)
- Meyer 9621, Pearl Village Hotel: footballfield on road to resort, Phuket, 10 m (BKF)
- Meyer 9622, Pearl Village Hotel: fallow field just outside entrance, Phuket, 20 m (BKF)
- Nielsen 685, Koh Chang, Trat (C)
- Nielsen 736, Koh Chang, Trat (C)
- Nielsen 738, Koh Chang, Trat (C)
- Niyomdham 84, Doi Phui, Mae Jam, Chiang Mai, 1600 m (BKF, C)
- Niyomdham 1561, Kok Mai Ruea, Tak Bai, Narathiwat, 0 m (AAU, E, K, BKF, C)
- Niyomdham 1646, Ku chum, Tak Bai, Narathiwat, 0 m (E, K, BKF, )
- Phenghklai 6999, Doi Inthanon National Park, Chiang Mai, 1300 m (BKF, C)
- Put 224, Baw Re, Chantaburi (K)
- Put 4317, Lat Bua Kao, Khorat, Nakhon Nayok (C, L)
- Sangkhachand 2, Lom kao, Nam Nao National Park, Phetchabun, 1000 m (BKF, C)
- Sangkhachand 142, Makham, Ban Arng, Chantaburi, 120 m (BKF, C)
- Sangkhachand 1040, Nai Chong, Krabi (BKF, C)
- Schmidt 27, Koh Chang, Trat (C)
- Schmidt 160, Koh Chang, Lan Dan, Trat (C)
- Schmidt 607 a, Koh Chang, Klong Majum, Trat (C)
- Seidenfaden 2049, Aerodrome Road, Bandon, Surat Thani, 6 m (SING, C)
- Seidenfaden 2184, Wat Sra Uhm. Bandon R., Surat Thani, 12 m (C)
- Seidenfaden 2339, Khun Tale Lake, Krat, Surat Thani (SING, C)
- Seidenfaden 2634, western slopes of Khao Sabab, Chantaburi, (C)
- Seidenfaden 2708, west of Khao Sabab, Chantaburi, 11 m (C)
- Shimizu 27226, Khao Pataew, Khao Pataew Wildlife Sanctuary, Thalang Distr., Phuket, 200 m (BKF, KYO, C)
- Singhasthit 407, Doi Suthep, Chiang Mai, 1100 m (BKF, C)
- Sørensen 448, east of Makam, Chantaburi, 100 m (C)
- Sørensen 647, Chawng, Forest Station west of Phattalung, Phattalung, 150 m (AAU, C)
- Sørensen 2617, Doi Suthep, Chiang Mai, 500 m (C)
- Sørensen 3215, Doi Suthep, Chiang Mai, 1700 m (C)
- Sørensen 7169, Koh Kut, Trat, 50 m (C)
- Suvarnasara 2, Khao Khieo, Nakhon Nayok, 1270 m (BKF, C)
USA
- Carlson 3782, Olaa to Black Sand Beach, Kalapana, Hawaii, 15 m (C)
- Schallert 18808, cultivated in my garden, Seminole Co.
- native of India, Florida (C)
VANUATU
- Curry 253, Aneitym Island
- Anelghowhat, Pinus caribea plantation west of Anelghowhat, 100 m (L)
VIETNAM
- Averjanov 168, Ba Mun, Quang Ninh (C)
- Averjanov 1152, Hon Than, Kien Giang (C)
- Feilberg s.n., Rismarker Nord for Saigon (C)
- Gaudichaud 111, Tourane (Da Nang), Quang Nam-Da Nang (P)
- Meyer 9613, Nationalroad QL 20, km 94, to Da Lat, Lam Dong, 300 m (MJG)
- Tsang 29011, Taai Wong Mo Shan, near Chuk-phai, Ha-coi, Tonkin, Tonkin (C)
COUNTRY UNKNOWN
- Kamphövener 1732 (C)
- Kamphövener 2516, Nanierory(?) (C)
- Koenig 1 (C)
- Koenig 2 (C)
- Koenig 3 (C)
- Koenig 4 (C)
- Liebmann s.n. (C)
- Liebmann s.n. (C)
- Miynt 1105, Penins. Ind. Orient. (C, GOET)
- Rinck s.n. (C)
- Vahl s.n. (C)
- Voigt s.n. (C)
- Wallich s.n., ex Ind. or (C).

- Protologue
- Synonyms
- Vernacular names
- Description
- Distribution, ecology and notes
- Collections

Melastoma malabathricum L. ssp. normale (D. Don) K. Meyer, Diss. Univ. Mainz (1999): 91. - Melastoma normale D. Don, Prodr. Fl. Nep. (1825): 220. - Type: Hamilton s.n., Nepal (K?, not seen).
Ramis undique setoso-pilosis, foliis ellipiticis acutis 5-nerviis basi rotundatis supra hispide pilosis subtus et lanuginosis, floribus ternis, calycibus squamulis lineari-setaceis ciliatis canis appressis omnino tectis.
Hab. in Nepalia. Hamilton
Obs. M. malabathricae affinis sed abunde distinctis.

- Melastoma wallichii DC., Prodr. 3 (1828): 146. - Type: Wallich 4039, prope Sikkim, India, 600 m (K!, P!, BM!).
- Melastoma houtteanum Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 291. - Type: van Houtte s.n., locality unknown (Burma?) (P!).
- Melastoma microphyllum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 293. - Type: Coll. ign. s.n., Inde ou Oceanie (P!).
- Melastoma velutinum Seem., Fl. Vit. (1865-73): 90. - Type: Armstrong 354, Port Essington, New Guinea (BM, not seen).
- Melastoma clarkeanum Cogn., Monogr. Phan. 7 (1891): 346. - Type: Helfer 2241, Tenasserim, Burma, 1000 m, 1837 (K!, P!).
- Melastoma osbeckioides Guillaumin, in Lecomte, Not. Syst. 2 (1909): 305. - Type: Thorel s.n., Cochonchine (P!).
- Melastoma eberhardtii Guillaumin, Fl. Gén. Indo-Chine 12 (1921): 889; Type: Eberhardt 2804, Chua-Thieu, haud cours du Bo-giang, Indochine (P!).
- Melastoma klossi Baker f., J. Nat. Hist. Soc. Siam 4 (1921): 132.
- Melastoma normale var. divergens Craib, Fl. Siam. 1 (1931): 682. - Type: Kerr 5540, Chiengdao, Muang Kawng, Chiang Mai, Thailand, 1400 m, 1 Jun 1921 (BK!, K!, BM!).
- Melastoma pubescens Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 87. - Type: Coll. ign. (Herb. Hasskarl) s.n., locality unknown (L!)

Vernacular names: Cay Mua (Vietnam), En-ah, I deng, Klong klang (Thailand), I si (Mussô, Thailand).

Shrubs 1.5--2.5 m high or small trees up to 4 m high; young branches quadrangular, covered with spreading or erect scales and bristles, bark brown; old branches terete, glabrescent. Leaves lanceolate, 6--12(--20) x 3--6(--8) cm, base rounded or acute, apex acuminate; lamina above strigose to pilose, beneath pilose; nerves 5 of 7, beneath covered with long spreading bristles or scales; petiole 7--25 mm long. Inflorescences terminal cymes. Flowers 5-merous; bracts lanceolate, 6--10 x 2--4 mm, outside hairy, inside glabrous. Hypanthium campanulate, 6--9 x 6--8 mm, covered with long appressed golden scales; sepals triangular-lanceolate, 6.5--8 x 3--4 mm, outside covered with appressed scales, inside glabrous; intersepalar emergences simple, c. 2 mm long. Petals obovate-cuneate, 22--27 x 17--20 mm, ciliate at the apex, violet. Stamens dimorphic; anthers of longer outer stamens c. 7.5 mm long, violet, connectives c. 5.5 mm long, ventrally curved, with 2 ventral appendages c. 1.5 mm long, filaments c. 8 mm long; anthers of shorter inner stamens c. 6.5 mm long, yellow, connectives not prolonged, ventrally with 2 appendages c. 1 mm long, filaments 7--8 mm long. Ovary slightly shorter than the hypanthium, crowned by bristles. Fruit a fleshy capsule, opening irregularly transversally at maturity, 7--10 x 7--10 mm, exposing the soft dark pulp with orange seeds.

Distribution - India (Karnataka, Sikkim), Bhutan, Nepal, Thailand (North, North-East), Laos, Vietnam, China, Malaysia (Perak), Indonesia (Sumatra, Java), New Caledonia, Philippines (Capiz).
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Ecology - Collections: 39. In disturbed evergreen, deciduous or coniferous forests, in savannas on sandstone plateaus, or on rocks at altitudes from 300--1900 m.
Notes - Melastoma malabathricum ssp. normale is a more northern taxon than its relatives, beeing mainly distributed from the edges of the Himalaya across northern Thailand to South China. Here it is the most common taxon of Melastoma. In tropical regions it is restricted to high mountains. The plants can be identified by their pilose leaves and brown scales on the branches.

BHUTAN
- Grierson 1355, Tama, lower Mangde Chu valley, 1100 m (K). CHINA. Rottboll s.n., East China (C)
- Tsang 28635, Chu Koo Nin Shue, Kwai Shan, Tsing-lo-kong village, Ho-yuen District, Guangdong (P)
- Tsui 311, Wan Tong Shan, Ying Tak Distr., Guangdong (SING)
INDIA
- Hooker s.n., Sikkim (C, GOET)
- Thomson s.n., Maisor & Carnatic, Karnataka (C, GOET))
INDONESIA
- Blume s.n., Java, Tjibiday, Jawa Barat (P)
- Nielsen 1285, Volcan Pintau, Sumatera Utara (C). LAOS. dOrléans s.n. (P). MALAYSIA. Everett FRI 13578, Taiping: 3.25 mile up road. Road verge hill side, over acidic rock, Perak, 680 m (SING, KEP)
NEPAL
- Barclay 2302, on trail Henje-Dampus, 1500 m (K)
- Grey-Wilson 266, Khare, east of Lumle, 1700 m (K)
- Wallich 4039, Napaulia (=Nepal) (K, BM)
- Wallich s.n., Napaulia (=Nepal) (C)
- Wallich s.n., Nepal (C)
NEW CALEDONIA
- Cheesman 3131, Tinchialit Camp, 600 m (BM)
- McKee 9752, Vallée de la Ouinné, 300 m (L)
PHILIPPINES
- Ramos BS 31277, Jamindan, Panay, Capiz (L)
THAILAND
- Beusekom 4799, Mae Kray, Phrae, 1000 m (BKF, C, K, P)
- Hansen 11156, Phu Miang, Phetchabun, 1200 m (BKF, C)
- Hansen 12787, Doi Khun Huai Pong, Chiang Mai, 1850 m (AAU, BKF, C, E, K, L, P)
- Hansen 12894, Doi Pae Poe, about 90 km northwest of Tak, Tak, 1400 m (AAU, BKF, C, E, K, L, P)
- Hennipman 3562, Udawn, Phu Luang, Loei, 1350 m (BKF, C, K, L, P)
- Kerr 5540, Chiengdao, Muang Kawng, Chiang Mai, 1400 m (BK, K, US photo, BM)
- Kerr 20062, Khao Krading, Loei, 1200 m (BK, C, K)
- Meyer s.n., Phu Rua Nationalpark: grasslands below summit, Loei, 1200 m (MJG)
- Nakkan 40, Phu Kradung, Samkae, Loei, 800 m (C)
- Smitinand 2612, Lom Kao, Phetchabun, 800 m (BKF, C)
- Sørensen 1561, Pah Hom Pak, Chiang Rai, 1400 m (C)
- Sørensen 2294, Phu Kradung, Loei, 1300 m (C)
- Sørensen 2734, Doi Suthep-Pui National Park: Doi Suthep, Chiang Mai, 1000 m (C)
- Suvarnakoses 1425, Sithan, Loei (C, BKF)
VIETNAM
- Bon 4287, Doi Ngang (P)
- Eberhardt 1428, Huê eh eur., Thùa Thiên Huê (P)
- Eberhardt 4017, Guyen Quang, Guyen Quang (P, HM)
- Petelot 898, Cho Ganh (P)
- Petelot s.n., Phu Tho, Vinh Phu (C)
- Simond s.n., Long-Tchéou, Tonkin (P)
COUNTRY UNKNOWN
- Coll. ign. s.n., Inde ou Oceanie (P).

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