Melastoma sanguineum Sims
Key to the varieties
1a. Petioles covered with long bristles 2--9 mm long a. var. sanguineum b. Petioles covered with appressed scales or glabrous 2. 2a. Hypanthium sparsely covered with short, broad, spreading scales c. 1.1--3.0 x 0.7--1.0 mm c. var. ranauense b. Hypanthium sparsely covered with small appressed scales 0.8--2.2 mm long b. var. laevifolium
7a. Melastoma sanguineum Sims var. sanguineum Sims
Plate legend: A. habit; B. larger outer and smaller inner stamen; C. petal; D. intersepalar emergence; E. fruit; F. longitudinal section through the hypanthium.
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Table of contents
1. Protologue
Melastoma sanguineum Sims, Bot. Mag. 48 (1821): t. 2241.
2. Synonyms
- Melastoma erectum Jack, Trans. Linn. Soc. London, Bot. 14 (1823): 5.
- Melastoma decemfidum Roxb. ex Jack, Trans. Linn. Soc. London, Bot. 14 (1823): 6. - Melastoma muricatum Hunter ex Ridl., J. As. Soc. Straits 53 (1909): 91. - Type: Wallich 4042, Penang, Malaysia (K!).
- Melastoma macrocarpum D. Don, Mem. Wern. Nat. Hist. Soc. 4 (1823): 289. - Type: Bot. Mag. (1801): t. 529.
- Melastoma dodecandrum Roxb. non Lour., Fl. India 2 (1832): 405.
- Melastoma porphyreum Zipp. ex Blume, Flora 14 (1831): 487. - Type: Blume s.n., Moluccas, Indonesia (K!, L!).
- Melastoma pulcherrimum Korth., in Temminck, Verhand. Nat. Gesch., Bot., (1839-1842): 231. - Type: Korthals s.n., Borneo, Indonesia (K!, L!).
- Melastoma boryanum Korth., Verh. Nat. Gesch. Ned. Bezitt., Bot. 7 (1844): 232. - Type: Korthals s.n., Borneo, Indonesia (K!, L!).
- Melastoma nitidum Korth., Verh. Nat. Gesch. Ned. Bezitt., Bot. 7 (1844): 234. - Type: Korthals s.n., G. Sakoembang, Borneo, Indonesia (K!).
- Melastoma zollingeri Naudin, Ann. Sci. Nat. Bot., Ser. 3, 13 (1850): 277. - Type: Zollinger 2495, Java, Indonesia (P, not seen).
- Melastoma gaudichaudianum Naudin, Ann. Sci. Nat., Ser. 3, 13 (1850): 278. - Type: Gaudichaud 110, Tourane, Vietnam, Jan 1837 (P!).
- Melastoma esquirolii H. Lév., Feddes Repert. Spec. Nov. Regni Veg. 8 (1910): 61. - Types: Léveillé s.n., Long-Ky, Yunnan, China,700 m, 22 Mar (P); Esquirol 873, Montagnes Hoang-Tsao-Pa, Kouy-Tchéou, China, Jun 1906 (P!); Esquirol 1538, Montagnes Hoang-Tsao-Pa, Kouy-Tchéou, China, 1909 (P!).
- Melastoma schizocarpum Ridl., J. Straits Branch Roy. As. Soc. 59 (1911): 104. - Type: Ridley 15095, Ginting Kaleok, Perlis, Malaysia, Mar 1910 (K!).
- Melastoma chevalieri Guillaumin, Bull. Soc. Bot. France 68 (1921): 3. - Type: Chevalier 40395, Vietnam. Langbian: Massif du Langbian, Dran. 1200 m, 29 Apr 1919 (P!).
- Melastoma horridum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 63. - Type: Bünnemeyer 11596, G. Bonthain, Celebes, Indonesia, 1350 m (L!).
- Melastoma bancanum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 74. - Type: Bünnemeyer 1906, P. Pinang, Teroe, Bangka, Sumatera Selatan, Indonesia, 80 m, 14 Nov 1917 (L!).
- Melastoma balinense Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 75. - Type: Sarip 409, G. Batoe kase, Bali, Indonesia, 1650 m, 14 Oct 1918 (L!).
- Melastoma vile Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 88. - Type: Bünnemeier 11765, Celebes, Indonesia, 1200 m (L!).
- Melastoma koordersii Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 94. - Type: Koorders 19865, Idjen Plateau, Jawa Timur, Indonesia, 1200 m (L!).
- Melastoma elbertii Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 96. - Type: Elbert 984, Lombok, Rindjani Gebirge, N Seite; unterhalb Tengengeah, Nusa Tenggara Barat, Indonesia, 1350 m, 3 May 1909 (L!).
- Melastoma curvisepalum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 101. - Type: Elbert 859, Rindjani-Vulkangebirge, N-Seite, Sartjang, Lombok, Nusa Tenggara Barat, Indonesia, 900 m, 2 May 1909 (L!).
- Melastoma curvisepalum var. crinitum Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 102; Type: Elbert 858, Sadjang, G. Rindjani, , Lombok, Nusa Tenggara Barat, Indonesia, 300-500 m (L!).
- Melastoma dendrisetosum C. Chen, J. South China Agricult. Univ. 4 (1983): 35. - Type: Hainan Expedition 89, China, Hainan: Lehui (SCBI, not seen).
- Melastoma sanguineum var. latisepalum C. Chen, J. South China Agricult. Univ. 4 (1983): 36. - Type: Chun s.n., China, Hainan: Dongfang (KUN, not seen).
3. Vernacular names
No vernacular names known!
4. Description
Shrubs or small trees up to 10 m high; young branches quadrangular, covered with appressed or spreading, red to brown scales or bristles; bark brown or grey. Leaves lanceolate or elliptic, seldom ovate, 2.8--17.4 x 1.0--5.8 cm, base rounded or acute, apex acute or acuminate; lamina glabrous or strigose, or seldom subvillous underneath, pale green or tinged red beneath; nerves 5 or 7, the marginal nerves unconspicious, covered with appressed or semi-erect scales or bristles, seldom glabrous, nerves often red; petiole 3--41 mm long, covered with erect or flexuose red bristles, 5--9 mm long. Inflorescences terminal cymes of 3--5 flowers, or flowers solitary. Flowers 5-merous, oocasionally 6- or 7-merous. Bracts ovate to lanceolate, 4--12 x 2--10 mm, the outside covered with bristles, the inside glabrous. Hypanthium 7--15 x 5--14 mm; covered with spreading, reddish bristles, 2--9 mm long, bristles deflexing with age; sepals deciduous, lanceolate or triangular, 5--17 x 1--5 mm, the outside covered with bristles, the inside glabrous or covered at the tip with appressed bristles; intersepalar emergences ca. 4--5 mm long, crowned by 4--5 mm long bristles. Petals cuneate-obovate, 33--46 x 22--32 mm, at the apical margin ciliate, violet. Stamens dimorphic. Anthers of longer outer stamens 12--18 mm long, slightly ventrally curved, violet; connectives 10--16 mm long, ventrally curved with 2 yellow, ventral lobes c. 1--3 mm long; filaments 8--12 mm long, yellow. Anthers of shorter stamens 8--14 mm long, ± straight, yellow, connective not prolonged but basal with 2 ventral appendages c. 1--2 mm long; filaments 5--8 mm long, yellow. Ovary nearly as long as the hypanthium, crowned by golden bristles. Style c. 37 mm long, ± straight. Fruits campanulate, fleshy capsules, 8--19 x 8--18 mm, rupturing irregularly longitudinally at maturity, exposing the solid, yellow pulp with orange seeds. Seeds minute, cochleate.
5. Distribution, ecology and notes
Distribution - Burma, Thailand, Malaysian Pensinsula, Sumatra, Borneo, Moluccas, South China, Indochina.
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Ecology - Collections: 87. In disturbed forests, along streams and roads, in open places and savannas at elevations of up to 2300 m.
Notes - Melastoma sanguineum var. sanguineum is a widespread species with considerable morphological variation especially in the size of the hypanthium emergences, but its vegetative characters are remarkably invariant.
6. Collections
BRUNEI
- Anderson 83/12, Bkt. Rerak, north side, Temburong, 1200 m (E)
- Coode 6571, Temburong river, just upstream from Wong Nguan rapids, Temburong, 120 m (C, K)
BURMA
- Lace 4632, Panigkyn Is Tale, Daune Range, Thalon, 1200 m (K)
CHINA
- Didrichsen 3217, Hongkong (C)
- Feilberg s.n., Happy Valley, Hongkong (C)
INDONESIA
- Blume s.n., Moluccas, Maluku (K, L)
- Elbert 4300, Flores
- Landschaft Endeh O., Doaria, Nusa Tenggara Timur, 500 m (L)
- Hansen 1208, Nangabulik, camp at logging road c. 25 km west of the village Sungai Buluh, Kalimantan Tengah, 300 m (C)
- Hansen 1293, Banjarmasin, hill with the ruins of the former Dutch governor’s residence close to Riamkanan, Kalimantan Selatan, 300 m (C)
- Korthals s.n., Borneo (K, L)
- Korthals s.n., Borneo (C)
- Korthals s.n., Borneo, Dausau (C)
- Korthals s.n., Borneo (C)
- Kostermans 191, Ndeki, west Flores, Nusa Tenggara Timur, 350 m (L)
- Kostermans 428, Lobok-besar, Bangka, Isl. Bangka, Sumatera Selatan, 20 m (P, L)
- Ploem s.n., Java (L)
- Verheijen 2518, Flores, Nusa Tenggara Timur (L)
- Voogd 1839, Bali (L)
MALAYSIA
- Ag. Hassan Chuchu SAN 62314, G. Silam, Lahad Datu Distr., Sabah, 750 m (SAN)
- Amin SAN 116344, Bayan, Ranau Distr., Sabah (SAN)
- Amin SAN 119811, Tompurongon, Ranau Distr., Sabah, (SAN)
- Awa S 47916, G. Pagon, Limbang, 5th Div., Sarawak, 1500 m (KEP, SAR, K, L, AAU, C)
- Awa S 47944, Bt. Pagon: Summit of Bt. Pagon, Limbang, 5th Division, Sarawak, 1770 m (SAR, AAU, C, L, K, KEP)
- Awa S 50669, Sg. Pa Mario, Ulu Limbang, route to Batu Lawi, Bario, 4th Division, Sarawak, 1530 m (SAR, KEP, C, L, K)
- Banyeng S 45090, Bt. Sadok, summit, Sarawak, (SAR, AAU, C, K)
- Burkill 3345, G. Jerai (Kedah Peak), Kedah, 960 m (SING, SAR, SAN)
- Chew RSNB 4404, Sg. Bembangan, Sabah, 1500 m (SAN, K, L)
- Clausing 166, G. santubong, at the summit, Sarawak, 810 m (UNIMAS)
- Clausing 202, near golf course, Mt. Penrissen, Sarawak (MJG)
- Clausing 203, near golf course, Mt. Penrissen, Sarawak (MJG) Clausing 218, Pig Hill, Mt. Kinabalu National Park, Sabah (Mt. Kinabalu Herbarium)
- Cockburn SAN 83348, near draining G. Lotung Inarat, Lamag Distr., Sabah, 390 m (SAR, SAN, K, L, KEP)
- Curtis 683, Penang: Government Hill, Penang, 400 m (K)
- Davis 69403, Kedah Peak: north side, Kedah, 1200 m (E)
- Fidilis SAN 106897, Sg. Saburan, Sepulut, Keningau Distr., Sabah, (SAN)
- Fidilis SAN 125637, Ulu Sg. Sembuan, Tulio area
- Keningau Distr., Sabah (SAN)
- Geh 201, Punan Busang, Belaga, 7th Div., Sarawak (SAR, SING, C)
- Hansen 1061, G. Lesung, Lingga, 2nd Div., Sarawak, 540 m (C)
- Hardial 227, Ayer Hitam Dam Reserve, Penang (C)
- Ilias S 26399, G. Murut: Path to G. Murut (Camp 4), Lawas, Sarawak, 2100 m (SAR, SING, SAN, K, L)
- J.C. 1594, Ulu Gombok: 17 mile south, Selangor (SING, K, L)
- Jack 54, Pulau Penang, Penang (BM)
- Kiah bin Hdj. Mohd. Salleh SK 409, Penang Hill, Penang, 900 m (SING)
- Kochummen FRI 18063, G. Jerai: Kedah Peak roadside near rimba reareasi, Kedah (SAN, KEP, L)
- Lee S 39993, Sg. Nawai, Linau Balui, Belaga, 7th Div., Sarawak, 800 m (KEP, SAR, UA, K, L, E)
- Madani SAN 81702, mile 87.5 Telupid, Hap Seng logging area
- Sandakan Distr., Sabah (SAN, KEP)
- Madani SAN 89371, Bt. Ampuan, Ranau Distr., Sabah (SAN)
- Maikin SAN 130704, Bidu-Bidu FR, Beluran Distr., Sabah (SAN)
- Mantor SAN 114071, Sg. Seliwan, Sepulut Distr., Sabah (SAN)
- Mantor SAN 118772, Sg. Millian, Nabawan Distr., Sabah (SAN)
- Meyer 9555, Penang: Penang Hill, Jalan Waterfall, Penang, 700 m (KEP)
- Meyer 9556, Penang: Penang Hill, Jalan Waterfall, Penang, 500 m (KEP)
- Meyer 9557, Penang: Penang Hill, Moniol Road above Botanical Garden, Penang, 100 m (KEP)
- Othman S 48934, Bt. Tebunan, Ulu Trusan, Lawas, 5th Division, Sarawak, 1250 m (SAR, L, K, KEP, AAU, C)
- Ridley 15095, Ginting Kaleok, Perlis (K)
- Shah 3648, Water catchment area at damp. Mt Ophir, Johore, 400 m (SING, KEP, C)
- Shukor 51, Kedah, 960 m (C)
- Sigin SAN 99702, Sg. Meliau, Beluran Distr., Sabah (SAN, PSU)
- Tong S 32939, Ulu Sg. Masia, Kota Forest Reserve, Lawas, 5th Division, Sarawak, 1400 m (SAR, K, L, SING, SAN, KEP)
- Wallich 4042, Penang, Penang (K)
- Whitmore FRI 12516, Central Kelantan, G. Stong eastern face, Kelantan (SING, KEP, L)
- Wong 2195, Tongod, G. Tingkar, Sabah, 150 m (SAN)
- Yii S 44459, Gunung Murud National Park, 4th Division, Sarawak, 2300 m (SAR, AAU, C)
THAILAND
- Bunchuai 1653, Non Sang, Udon Thani (BKF, C, L)
- Bunpheng 994, Phu Luang, Loei (BKF, C)
- Geesink 5171, Khao Pawta Luang Kheo, Phangnga, 900 m (AAU, C, E, L)
- Iwatsuki T 7481, Nakhon Nayok: Khao Kiew, a high ridge in Khao Yai National Park, Prachinburi, 1250 m (C)
- Larsen 43239, Ngao waterfall south of Ranong, Ranong, 150 m (AAU, PSU)
- Meyer 955, Hat Yai: Prince of Songkhla University, Kuan Mod Dang Hill, in front of the guest house, Songkhla (BKF)
- Meyer 957, Hat Yai: Ko Hong Hill, near summit, Songkhla, 350 m (BKF)
- Meyer 958, Hat Yai: Ko Hong Hill, road to summit, Songkhla, 200 m (BKF)
- Meyer 9510, Ton Nga Chang Wildlife Sanctuary: Ton Nga Chang waterfalls, Songkhla, 200 m (BKF)
- Meyer 9511, Ton Nga Chang Wildlife Sanctuary: Ton Nga Chang waterfalls, Songkhla, 200 m (BKF)
- Meyer 9512, Ton Nga Chang Wildlife Sanctuary: Ton Nga Chang waterfalls, Songkhla, 200 m (BKF)
- Meyer 9517, Hat Yai: Prince of Songkhla University, Kuan Mod Dang Hill, in front of the guest house, Songkhla (BKF)
- Meyer s.n., Phu Rua Nationalpark: stream near camp-site, Loei, 1000 m (MJG)
- Niyomdham 1560, Tak Bai, Kok Mai Ruen, Narathiwat, 0 m (AAU, BKF, E, K, C, L)
- Schmidt 607a, Koh Chang, Nipple, Trat, 600 m (C)
- Schmidt 672a, Koh Chang, Nipple, Trat, 600 m (C)
- Sørensen 3217, Doi Suthep-Pui National Park: Doi Suthep, Chiang Mai, 1700 m (C)
VIETNAM
- Bon 5588, Cua Bang (C)
- Meyer 9614, Bao Loc, Dam Bri Waterfall, area behind gate, Lam Dong, 800 m (HNU)
- Petelot 952, Cho Ganh, Ha Nam Ninh (C, P)
- Poilane 4566, massif de Cô Inh près de Nha Trang, Nha Trang, 400 m (C, P, HM)
- Tsang 29243, Taai Wong Mo Shan, near Chuk-phai Ha-coi, Tonkin (SING, C, P)
- Tsang 30060, Sai Wong Mo Shan (Sai Vong Mo Leng), Lung Wan Village, Dam-ha, Tonkin (SING, C, P)
- Tsang 39212, Taai Wong Mo Shan, near Chuk-phai, Ha-coi, Tonkin (C).
7b. Melastoma sanguineum Sims var. laevifolium (Merr.) K. Meyer
Table of contents
1. Protologue
Melastoma sanguineum Sims var. laevifolium (D. Don) K. Meyer, Diss. Univ. Mainz (1999): 112. - Melastoma laevifolium Merr., J. Straits Branch Roy. Asiat. Soc. 86 (1922): 337. - Type: Ramos BS 1132, Sandakan and vicinity, Sabah, Malaysia, Dec 1920 (K!).
Frutex erectus, circiter 2 m altus, obscure et parcissime strigillosus; ramis ramulisque tenuibus, teretibus, ramis glabris, ramulis rubro-brunneis, paleis adpressis, minutis sparsis obscuris instructis vel sublaevibus; foliis lanceolatis, coriaceis, rigidis, 4-9 cm longis, acuminatis, basi acutis, 3-nerviis, supra olivaceis, laevibus, parcissime strigillosis, subtus viridibus, sublaevibus, paleis minutis paucis instructis; floribus terminalibus, solitariis, 5-meris, calyci obscure parceque paleaceo, paleis ovatis acutis adpressis vix 0.5 mm longis instructo, lobis latis, brevibus, circiter 2 mm longis; antherarum majorum connectivo producto.
An erect shrub about 2 m high nearly glabrous throughout, the branches and branchlets terete, smooth, the ultimate branchlets reddish-brown, 1.5 mm in diameter or less, supplied with very few widely scattered, appressed, ovate scales about 0.2 mm long. Leaves lanceolate, coriaceus, rigid, 4 to 9 cm long, 1 to 1.5 cm wide, narrowed upward to the acuminate apex, the base acute, 3-nerved, the upper surface olivaceus, somewhat shining smooth, obscurely and very sparsely strigulose, the lower surface paler and with few widely scattered, minute, appressed scales similar to those on the branchlets, the midrib, nerves and reticulations reddish in contrast with the greenish surface; petioles slender, 7 to 10 mm long, with very few minute, appressed scales. Flowers terminal, solitary, 5-merous, their pedicels about 1 cm long. Calyx about 1 cm long, subcylindric, acute at the base, about 8 mm in diameter, the lobes short, broad, subobtuse, about 2 mm in lenght, the pedicels, tube, and lobes supplied with minute, apressed, widely scattered, triangular-ovate scales 0.5 mm long or less. Petals oblong-obovate, about 3 cm long, 1.5 cm wide, narrowed below to the subacute base, the apex acute or somewhat obtuse. Stamens unequal, the filaments of the shorter ones 9 mm long, their anthers curved and about as long as the filaments, the longer filaments 12 mm long, the connective produced about 14 mm, 2-appendiculate, the appendages stout, 2 mm long. Top of the ovary produced, about 2 mm long around the base of the style and conspicuously ciliate.
British North Borneo, Sandakan, Ramos 1132, September, 1920. In thickets near the seashore.
A species perhaps most closely allied to Melastoma nitidum Zoll.; differing, however, in numerous characters. In the genus it is well characterized by being nearly smooth and glabrous, the scales on the branchlets, petioles, pedicels, and leaves being inconspicuous, minute, and widely scattered, the upper surface of the leaves being entire glabrous.
2. Synonyms
- Melastoma decipiens Bakh. f., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 91 (1943): 85. - Type: Sarip 51, G. Kelatakan, Bali, Indonesia, 585 m (L!).
3. Vernacular names
Vernacular names: Kemalahidi (Borneo), Ahe rápéss (Sumba), Sose bereq (Timor).
4. Description
Shrubs up to 2 m high; young branches quadrangular, covered with appressed, red to brown scales; bark dark brown. Leaves lanceolate, 9--11.5 x 1.5--3 cm, base acute, apex acute or acuminate; surfaces glabrous or strigose; nerves 5 or 7, sparsely covered with small appressed scales; petiole 5--25 mm long, covered with small appressed scales. Inflorescences terminal cymes of 3--5 flowers. Flowers 5-merous; bracts deciduous, not seen. Hypanthium 13--15 x 8--12 mm; sparsely covered with small appressed scales c. 0.8--2.2 mm long; sepals deciduous, lanceolate, 15--22 x 3--7 mm; intersepalar emergences subulate, ca. 4--5 mm long Petals not seen, violet. Stamens not seen. Ovary nearly as long as the hypanthium, crowned by golden bristles. Style c. 37 mm long. Fruits campanulate, fleshy capsules, 13--17 x 9--15mm.
5. Distribution, ecology and notes
Distribution - Borneo, Lesser Sunda Islands, Timor.
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Ecology - Collections: 7. In disturbed or secondary lowland rain forests.
Notes - This variety differs from var. sanguineum by its very small appressed scales. The examination of many specimens reveals continuous variation in bristle length and -width between the type variant and the specimens cited
6. Collections
INDONESIA
- Balgooy 5376, G. Palung Nature Reserve, c. 100 km south of Pontianak, Kalimantan Barat, 600 m (L)
- Friedberg 287, Lolotoi, Timor Timur (Portugal) (L)
- Iboet 558, Sumba, Kananggar, Nusa Tenggara Timur (L)
- Jaag 943, Pisigomo, Lantoha, Alor, Nusa Tenggara Timur, 650 m (L)
MALAYSIA
- Meijer SAN 131705, G. Walker
- Sandakan Distr., Sabah, 229 m (SAN)
- Ramos BS 1132, Sandakan and vicinity, Sabah (K, P, L)
- WM SAN 20152, Buli-Sim-Sim, Sandakan, Sabah, 60 m (L).
7c. Melastoma sanguineum Sims var. ranauense K. Meyer
Table of contents
1. Protologue
Melastoma sanguineum var. ranauensis K. Meyer, Diss. Univ. Mainz (1999): 113. - Holotype: Amin SAN 116344, Bayan, Ranau Distr., Sabah, Malaysia (SAN!).
Melastoma sanguineum var. ranauensis a var. sanguineum hypanthii squamae brevae sparse patentibus distinguenda.
2. Synonyms
No synonyms known!
3. Vernacular names
No vernacular names known!
4. Description
Shrubs or small trees up to 3 m high; young branches quadrangular, covered with appressed, red to brown scales; bark grey or brown; older branches terete, glabrescent. Leaves lanceolate, 5.5--11.5 x 2--3 cm, base and apex acute; lamina strigose; nerves 5, sparsely covered with appressed scales; petiole 5--20 mm long, covered with small appressed scales. Inflorescences terminal few-flowered cymes or flowers solitary. Flowers 5-merous; bracts deciduous, not seen. Hypanthium sparsely covered with short spreading scales c. 1.1--3.0 x 0.7--1.0 mm, sepals deciduous, lanceolate, 15--22 x 3--7 mm; intersepalar emergences subulate, ca. 4--5 mm long. Petals not seen, notably violet. Stamens not seen. Ovary slightly longer than the hypanthium, crowned by golden bristles. Fruits campanulate, fleshy capsules, 11--15 x 10--12 mm.
5. Distribution, ecology and notes
Distribution - Malaysia: Sabah, in the vicinity of Ranau.
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Ecology - Collections: 4. Except of the annotation ”habitat: hillside” nothing is known about the ecology of this variety.
Notes - This variety is known only from some collections around Ranau near the Mount Kinabalu National Park, Sabah, Malaysia. It is easily recognized by its spreading hypanthium scales, whereas the leaves are similar to those of the type variety.
6. Collections
MALAYSIA
- Amin SAN 116344, Bayan, Ranau Distr., Sabah (SAN)
- Amin SAN 119811, Tompurongon, Ranau Distr., Sabah (SAN)
- Chew RSNB 4404, Sg. Bembangan, Sabah, 1500 m (SAN, K, L)
- Madani SAN 89371, Bt. Ampuan, Ranau Distr., Sabah (SAN).
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